FORM 4 [ ] Check this box if no longer subject to Section 16. Form 4 or Form 5 obligations may continue. See Instruction 1(b).         
UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549

STATEMENT OF CHANGES IN BENEFICIAL OWNERSHIP OF SECURITIES
                                                                                  
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Filed pursuant to Section 16(a) of the Securities Exchange Act of 1934 or Section 30(h) of the Investment Company Act of 1940
                      

1. Name and Address of Reporting Person *

CAVANAGH RICHARD
2. Issuer Name and Ticker or Trading Symbol

BLACKROCK MUNIHOLDINGS FUND, INC. [ MHD ]
5. Relationship of Reporting Person(s) to Issuer (Check all applicable)

__X__ Director                    _____ 10% Owner
_____ Officer (give title below)    _____ Other (specify below)
(Last)          (First)          (Middle)

55 EAST 52ND STREET
3. Date of Earliest Transaction (MM/DD/YYYY)

3/8/2021
(Street)

NEW YORK, NY 10055
(City)        (State)        (Zip)
4. If Amendment, Date Original Filed (MM/DD/YYYY)

 
6. Individual or Joint/Group Filing (Check Applicable Line)

_X _ Form filed by One Reporting Person
___ Form filed by More than One Reporting Person

Table I - Non-Derivative Securities Acquired, Disposed of, or Beneficially Owned
1.Title of Security
(Instr. 3)
2. Trans. Date 2A. Deemed Execution Date, if any 3. Trans. Code
(Instr. 8)
4. Securities Acquired (A) or Disposed of (D)
(Instr. 3, 4 and 5)
5. Amount of Securities Beneficially Owned Following Reported Transaction(s)
(Instr. 3 and 4)
6. Ownership Form: Direct (D) or Indirect (I) (Instr. 4) 7. Nature of Indirect Beneficial Ownership (Instr. 4)
Code V Amount (A) or (D) Price
Common Stock 3/8/2021  J(1)  323 A (1)443 D  
Common Stock 3/8/2021  J(2)  115 A (2)558 D  
Common Stock 3/8/2021  J(3)  112 A (3)670 D  
Common Stock 3/8/2021  J(4)  97 A (4)767 D  

Table II - Derivative Securities Beneficially Owned (e.g., puts, calls, warrants, options, convertible securities)
1. Title of Derivate Security
(Instr. 3)
2. Conversion or Exercise Price of Derivative Security3. Trans. Date3A. Deemed Execution Date, if any4. Trans. Code
(Instr. 8)
5. Number of Derivative Securities Acquired (A) or Disposed of (D)
(Instr. 3, 4 and 5)
6. Date Exercisable and Expiration Date7. Title and Amount of Securities Underlying Derivative Security
(Instr. 3 and 4)
8. Price of Derivative Security
(Instr. 5)
9. Number of derivative Securities Beneficially Owned Following Reported Transaction(s) (Instr. 4)10. Ownership Form of Derivative Security: Direct (D) or Indirect (I) (Instr. 4)11. Nature of Indirect Beneficial Ownership (Instr. 4)
CodeV(A)(D)Date ExercisableExpiration DateTitleAmount or Number of Shares

Explanation of Responses:
(1) Effective as of March 8, 2021, BlackRock Municipal Income Investment Quality Trust (the "Target Fund") was reorganized into BlackRock MuniHoldings Fund, Inc. (the "Acquiring Fund"). In the Reorganization, common shareholders of the Target Fund received common shares of the Acquiring Fund having a value equal to the aggregate NAV of the Target Fund common shares surrendered as determined at the close of business on March 5, 2021, less the costs of the Reorganization. As of March 5, 2021, the Target Fund reported a NAV per share of $15.2503 and the Acquiring Fund reported a NAV per share of $16.9927. The conversion ratio for the Target Fund's common shares was 0.89746185. In the Reorganization, Richard E. Cavanagh received 323 common shares (and cash for fractional shares, if any) of the Acquiring Fund in exchange for his 360 common shares of the Target Fund.
(2) Effective as of March 8, 2021, BlackRock Municipal Bond Trust (the "Target Fund") was reorganized into BlackRock MuniHoldings Fund, Inc. (the "Acquiring Fund"). In the Reorganization, common shareholders of the Target Fund received common shares of the Acquiring Fund having a value equal to the aggregate NAV of the Target Fund common shares surrendered as determined at the close of business on March 5, 2021, less the costs of the Reorganization. As of March 5, 2021, the Target Fund reported a NAV per share of $16.4314 and the Acquiring Fund reported a NAV per share of $16.9927. The conversion ratio for the Target Fund's common shares was 0.96696817. In the Reorganization, Richard E. Cavanagh received 115 common shares (and cash for fractional shares, if any) of the Acquiring Fund in exchange for his 119 common shares of the Target Fund.
(3) Effective as of March 8, 2021, BlackRock MuniHoldings Fund II, Inc. (the "Target Fund") was reorganized into BlackRock MuniHoldings Fund, Inc. (the "Acquiring Fund"). In the Reorganization, common shareholders of the Target Fund received common shares of the Acquiring Fund having a value equal to the aggregate NAV of the Target Fund common shares surrendered as determined at the close of business on March 5, 2021, less the costs of the Reorganization. As of March 5, 2021, the Target Fund reported a NAV per share of $15.7316 and the Acquiring Fund reported a NAV per share of $16.9927. The conversion ratio for the Target Fund's common shares was 0.92578578. In the Reorganization, Richard E. Cavanagh received 112 common shares (and cash for fractional shares, if any) of the Acquiring Fund in exchange for his 121 common shares of the Target Fund.
(4) Effective as of March 8, 2021, BlackRock MuniHoldings Quality Fund, Inc. (the "Target Fund") was reorganized into BlackRock MuniHoldings Fund, Inc. (the "Acquiring Fund"). In the Reorganization, common shareholders of the Target Fund received common shares of the Acquiring Fund having a value equal to the aggregate NAV of the Target Fund common shares surrendered as determined at the close of business on March 5, 2021, less the costs of the Reorganization. As of March 5, 2021, the Target Fund reported a NAV per share of $13.8681 and the Acquiring Fund reported a NAV per share of $16.9927. The conversion ratio for the Target Fund's common shares was 0.81612104. In the Reorganization, Richard E. Cavanagh received 97 common shares (and cash for fractional shares, if any) of the Acquiring Fund in exchange for his 120 common shares of the Target Fund.

Reporting Owners
Reporting Owner Name / Address
Relationships
Director10% OwnerOfficerOther
CAVANAGH RICHARD
55 EAST 52ND STREET
NEW YORK, NY 10055
X



Signatures
/s/ Dean A. Caruvana as Attorney-in-Fact3/10/2021
**Signature of Reporting PersonDate

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