FORM 4
[ ] Check this box if no longer subject to Section 16. Form 4 or Form 5 obligations may continue. See Instruction 1(b).         
UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549

STATEMENT OF CHANGES IN BENEFICIAL OWNERSHIP OF SECURITIES
                                                                                  
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Filed pursuant to Section 16(a) of the Securities Exchange Act of 1934, Section 17(a) of the Public
Utility Holding Company Act of 1935 or Section 30(f) of the Investment Company Act of 1940
                      

1. Name and Address of Reporting Person *

Sullivan Timothy P
2. Issuer Name and Ticker or Trading Symbol

Ancestry.com Inc. [ ACOM ]
5. Relationship of Reporting Person(s) to Issuer (Check all applicable)

__ X __ Director                      _____ 10% Owner
__ X __ Officer (give title below)      _____ Other (specify below)
President and CEO
(Last)          (First)          (Middle)

360 WEST 4800 NORTH
3. Date of Earliest Transaction (MM/DD/YYYY)

11/18/2011
(Street)

PROVO, UT 84604
(City)        (State)        (Zip)
4. If Amendment, Date Original Filed (MM/DD/YYYY)

 
6. Individual or Joint/Group Filing (Check Applicable Line)

_ X _ Form filed by One Reporting Person
___ Form filed by More than One Reporting Person

Table I - Non-Derivative Securities Acquired, Disposed of, or Beneficially Owned
1.Title of Security
(Instr. 3)
2. Trans. Date 2A. Deemed Execution Date, if any 3. Trans. Code
(Instr. 8)
4. Securities Acquired (A) or Disposed of (D)
(Instr. 3, 4 and 5)
5. Amount of Securities Beneficially Owned Following Reported Transaction(s)
(Instr. 3 and 4)
6. Ownership Form: Direct (D) or Indirect (I) (Instr. 4) 7. Nature of Indirect Beneficial Ownership (Instr. 4)
Code V Amount (A) or (D) Price
Common Stock, $0.001 par value   11/18/2011     M    86956   A $4.6   411030   D    
Common Stock, $0.001 par value   11/18/2011     M    37036   A $5.4   448066   D    
Common Stock, $0.001 par value   11/21/2011     G    324074   D $ 0   123992   D    
Common Stock, $0.001 par value   11/21/2011     G    324074   A $ 0   324074   I   By Purefoy LLC  

Table II - Derivative Securities Beneficially Owned ( e.g. , puts, calls, warrants, options, convertible securities)
1. Title of Derivate Security
(Instr. 3)
2. Conversion or Exercise Price of Derivative Security 3. Trans. Date 3A. Deemed Execution Date, if any 4. Trans. Code
(Instr. 8)
5. Number of Derivative Securities Acquired (A) or Disposed of (D)
(Instr. 3, 4 and 5)
6. Date Exercisable and Expiration Date 7. Title and Amount of Securities Underlying Derivative Security
(Instr. 3 and 4)
8. Price of Derivative Security
(Instr. 5)
9. Number of derivative Securities Beneficially Owned Following Reported Transaction(s) (Instr. 4) 10. Ownership Form of Derivative Security: Direct (D) or Indirect (I) (Instr. 4) 11. Nature of Indirect Beneficial Ownership (Instr. 4)
Code V (A) (D) Date Exercisable Expiration Date Title Amount or Number of Shares
Common Stock Option (right to buy)   $4.6   11/18/2011     M         86956      (1) 11/15/2015   Common Stock, $0.001 par value   86956   $ 0   1649478   D    
Common Stock Option (right to buy)   $5.4   11/18/2011     M         37036      (2) 3/27/2018   Common Stock, $0.001 par value   37036   $ 0   334964   D    
Common Stock Option (right to buy)   $39.34                      (3) 5/16/2021   Common Stock, $0.001 par value   300000     300000   D    
Restricted Stock Units   $ 0   (4)                    (5)   (5) Common Stock, $0.001 par value   150000     150000   D    

Explanation of Responses:
( 1)  Option granted on November 15, 2005. This option, originally representing a right to purchase 2,000,000 shares, is fully vested and immediately exercisable.
( 2)  Option granted on March 27, 2008. This option, originally representing a right to purchase 372,000 shares, vests over four years with 1/4 of the total number of shares subject to the option having vested on January 1, 2009 and 1/48 of the total number of shares subject to the option vesting monthly thereafter.
( 3)  Option granted on May 16, 2011. 33% of the total number of shares subject to the option will vest on each of the third and fourth anniversaries of the date of grant and 34% of the total number of shares subject to the option will vest on the fifth anniversary of the date of grant.
( 4)  Each restricted stock unit represents a contingent right to receive one share of Ancestry.com Inc. common stock.
( 5)  Restricted stock units granted on May 16, 2011. 33% of the restricted stock units granted will vest on each of the third and fourth anniversaries of the date of the grant and 34% of the restricted stock units granted will vest on the fifth anniversary of the date of the grant. Upon vesting, the reporting person will receive a number of shares of common stock equal to the number of restricted stock units that have vested.

Reporting Owners
Reporting Owner Name / Address
Relationships
Director 10% Owner Officer Other
Sullivan Timothy P
360 WEST 4800 NORTH
PROVO, UT 84604
X
President and CEO

Signatures
/s/ Christopher Brookhart, Attorney-in-fact for Timothy Sullivan 11/22/2011
** Signature of Reporting Person Date


Reminder: Report on a separate line for each class of securities beneficially owned directly or indirectly.
* If the form is filed by more than one reporting person, see Instruction 4(b)(v).
** Intentional misstatements or omissions of facts constitute Federal Criminal Violations. See 18 U.S.C. 1001 and 15 U.S.C. 78ff(a).
Note: File three copies of this Form, one of which must be manually signed. If space is insufficient, see Instruction 6 for procedure.
Persons who respond to the collection of information contained in this form are not required to respond unless the form displays a currently valid OMB control number.
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