FORM 4
[ ] Check this box if no longer subject to Section 16. Form 4 or Form 5 obligations may continue. See Instruction 1(b).         
UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549

STATEMENT OF CHANGES IN BENEFICIAL OWNERSHIP OF SECURITIES
                                                                                  
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Filed pursuant to Section 16(a) of the Securities Exchange Act of 1934, Section 17(a) of the Public
Utility Holding Company Act of 1935 or Section 30(f) of the Investment Company Act of 1940
                      

1. Name and Address of Reporting Person *

PARKER VICTOR E JR 		2. Issuer Name and Ticker or Trading Symbol

Ancestry.com Inc. [ ACOM ] 		5. Relationship of Reporting Person(s) to Issuer (Check all applicable)

__ X __ Director                      _____ 10% Owner
_____ Officer (give title below)      _____ Other (specify below)
(Last)          (First)          (Middle)

360 WEST 4800 NORTH 		3. Date of Earliest Transaction (MM/DD/YYYY)

11/10/2009
(Street)

PROVO, UT 84604
(City)        (State)        (Zip)
4. If Amendment, Date Original Filed (MM/DD/YYYY)

 
6. Individual or Joint/Group Filing (Check Applicable Line)

_ X _ Form filed by One Reporting Person
___ Form filed by More than One Reporting Person

Table I - Non-Derivative Securities Acquired, Disposed of, or Beneficially Owned
1.Title of Security
(Instr. 3) 		2. Trans. Date 2A. Deemed Execution Date, if any 3. Trans. Code
(Instr. 8) 		4. Securities Acquired (A) or Disposed of (D)
(Instr. 3, 4 and 5) 		5. Amount of Securities Beneficially Owned Following Reported Transaction(s)
(Instr. 3 and 4) 		6. Ownership Form: Direct (D) or Indirect (I) (Instr. 4) 7. Nature of Indirect Beneficial Ownership (Instr. 4)
Code V Amount (A) or (D) Price
Common Stock, $0.001 par value   11/10/2009     S    2400444   D $13.50   22805668   I   (1) (2) By limited partnership (2)  

Table II - Derivative Securities Beneficially Owned ( e.g. , puts, calls, warrants, options, convertible securities)
1. Title of Derivate Security
(Instr. 3) 		2. Conversion or Exercise Price of Derivative Security 3. Trans. Date 3A. Deemed Execution Date, if any 4. Trans. Code
(Instr. 8) 		5. Number of Derivative Securities Acquired (A) or Disposed of (D)
(Instr. 3, 4 and 5) 		6. Date Exercisable and Expiration Date 7. Title and Amount of Securities Underlying Derivative Security
(Instr. 3 and 4) 		8. Price of Derivative Security
(Instr. 5) 		9. Number of derivative Securities Beneficially Owned Following Reported Transaction(s) (Instr. 4) 10. Ownership Form of Derivative Security: Direct (D) or Indirect (I) (Instr. 4) 11. Nature of Indirect Beneficial Ownership (Instr. 4)
Code V (A) (D) Date Exercisable Expiration Date Title Amount or Number of Shares

Explanation of Responses:
( 1)  Shares sold include (i) 1,755,277 shares of common stock owned by Spectrum Equity Investors V, L.P.; (ii) 8,289 shares of common stock owned by Spectrum V Investment Managers' Fund, L.P.; (iii) 626,352 shares of common stock owned by Spectrum Equity Investors III, L.P.; and (iv) 10,526 shares of common stock owned by Spectrum III Investment Managers' Fund, L.P. that the respective Spectrum entities sold as part of Ancestry.com Inc.'s initial public offering, which was completed on November 10, 2009.
( 2)  Shares of common stock held after sale consists of an aggregate of 22,805,668 shares, including (i) 16,676,204 shares held by Spectrum Equity Investors V, L.P.; (ii) 78,748 shares held by Spectrum V Investment Managers' Fund, L.P.; (iii) 5,950,719 shares held by Spectrum Equity Investors III, L.P.; and (iv) 99,997 shares held by Spectrum III Investment Managers' Fund, L.P. Mr. Parker is a managing director of the general partner of the general partner of Spectrum Equity Investors V, L.P. and a managing director of the general partner of Spectrum V Investment Managers' Fund, L.P. Mr. Parker is also a limited partner of the general partner of Spectrum Equity Investors V, L.P., a limited partner of Spectrum V Investment Managers' Fund, L.P., a limited partner of the general partner of Spectrum Equity Investors III, L.P. and a limited partner of Spectrum III Investment Managers' Fund, L.P. Mr. Parker disclaims beneficial ownership of these shares.

Reporting Owners
Reporting Owner Name / Address
Relationships
Director 10% Owner Officer Other
PARKER VICTOR E JR
360 WEST 4800 NORTH
PROVO, UT 84604 		X



Signatures
Christopher Brookhart, Attorney-in-Fact 11/10/2009
** Signature of Reporting Person Date


Reminder: Report on a separate line for each class of securities beneficially owned directly or indirectly.
* If the form is filed by more than one reporting person, see Instruction 4(b)(v).
** Intentional misstatements or omissions of facts constitute Federal Criminal Violations. See 18 U.S.C. 1001 and 15 U.S.C. 78ff(a).
Note: File three copies of this Form, one of which must be manually signed. If space is insufficient, see Instruction 6 for procedure.
Persons who respond to the collection of information contained in this form are not required to respond unless the form displays a currently valid OMB control number.
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