- Initial Statement of Beneficial Ownership (3)

FORM 3	UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 INITIAL STATEMENT OF BENEFICIAL OWNERSHIP OF SECURITIES	OMB APPROVAL OMB Number: 3235-0104 Expires: February 28, 2011 Estimated average burden hours per response... 0.5
	Filed pursuant to Section 16(a) of the Securities Exchange Act of 1934, Section 17(a) of the Public Utility Holding Company Act of 1935 or Section 30(h) of the Investment Company Act of 1940

1. Name and Address of Reporting Person ^*

Spectrum Equity Investors V L P

2. Date of Event Requiring Statement (MM/DD/YYYY) 11/4/2009

3. Issuer Name and Ticker or Trading Symbol

Ancestry.com Inc. [ACOM]

(Last) (First) (Middle)

333 MIDDLEFIELD ROAD, SUITE 200

4. Relationship of Reporting Person(s) to Issuer (Check all applicable)

_____ Director ___ X ___ 10% Owner
_____ Officer (give title below) _____ Other (specify below)

(Street)

MENLO PARK, CA 94025

(City) (State) (Zip)

5. If Amendment, Date Original Filed (MM/DD/YYYY)

6. Individual or Joint/Group Filing (Check Applicable Line)

___ Form filed by One Reporting Person
_ X _ Form filed by More than One Reporting Person

Table I - Non-Derivative Securities Beneficially Owned
1.Title of Security (Instr. 4)	2. Amount of Securities Beneficially Owned (Instr. 4)	3. Ownership Form: Direct (D) or Indirect (I) (Instr. 5)	4. Nature of Indirect Beneficial Ownership (Instr. 5)
Common Stock, $0.001 par value	25688319 ⁽¹⁾	D ⁽²⁾ ⁽³⁾ ⁽⁴⁾

*Table II - Derivative Securities Beneficially Owned ( e.g.* , puts, calls, warrants, options, convertible securities)**
1. Title of Derivate Security (Instr. 4)	2. Date Exercisable and Expiration Date (MM/DD/YYYY)		3. Title and Amount of Securities Underlying Derivative Security (Instr. 4)		4. Conversion or Exercise Price of Derivative Security	5. Ownership Form of Derivative Security: Direct (D) or Indirect (I) (Instr. 5)	6. Nature of Indirect Beneficial Ownership (Instr. 5)
1. Title of Derivate Security (Instr. 4)	Date Exercisable	Expiration Date	Title	Amount or Number of Shares	4. Conversion or Exercise Price of Derivative Security		6. Nature of Indirect Beneficial Ownership (Instr. 5)

Explanation of Responses:
( 1)	This Form 3 should be read in conjunction with the other Form 3 filed by Spectrum Equity Investors V, L.P. ("SEI V") as of the date hereof. Each of such Forms 3 reports the total number of shares of common stock owned by SEI V and its affiliates as described below. The amounts set forth in the two Forms 3 should not be added to reach a total.
( 2)	Consists of an aggregate of 25,688,319 shares, including 18,431,481 shares held by SEI V, the general partner of which is Spectrum Equity Associates V, L.P., the general partner of which is SEA V Management, LLC, over which Brion B. Applegate, William P. Collatos, Kevin J. Maroni, Randy J. Henderson, Michael J. Kennealy, Victor E. Parker, Benjamin M. Coughlin and Christopher T. Mitchell exercise voting and dispositive power; 87,037 shares held by Spectrum V Investment Managers' Fund, L.P. ("IMF V"), the general partner of which is SEA V Management, LLC, over which Brion B. Applegate, William P. Collatos, Kevin J. Maroni, Randy J. Henderson, Michael J. Kennealy, Victor E. Parker, Benjamin M. Coughlin and Christopher T. Mitchell exercise voting and dispositive power;
( 3)	6,577,071 shares held by Spectrum Equity Investors III, L.P. ("SEI III"), the general partner of which is Spectrum Equity Associates III, L.P., over which Brion B. Applegate, William P. Collatos, Kevin J. Maroni and Randy J. Henderson exercise voting and dispositive power; 461,426 shares held by SEI III Entrepreneurs' Fund, L.P. ("Entrepreneurs' III"), the general partner of which is SEI III Entrepreneurs' LLC, over which Brion B. Applegate, William P. Collatos, Kevin J. Maroni and Randy J. Henderson exercise voting and dispositive power; 110,523 shares held by Spectrum III Investment Managers' Fund, L.P. ("IMF III," and together with SEI V, IMF V, SEI III and Entrepreneurs' III, the "Spectrum Funds"), over which Brion B. Applegate, William P. Collatos, Kevin J. Maroni and Randy J. Henderson exercise voting and dispositive power and are the general partners, 12,793 shares held by Brion B. Applegate, 6,390 shares held by William P. Collatos and 1,598 shares held by Randy J. Henderson.
( 4)	Each of the controlling entities, individual general partners and managing directors of the Spectrum Funds, as the case may be, including Mr. Parker who is a managing director of the general partner of the general partner of SEI V and a managing director of the general partner of IMF V, and serves on Ancestry.com Inc.'s board of directors, Brion B. Applegate, William P. Collatos, Kevin J. Maroni, Randy J. Henderson, Michael J. Kennealy, Victor E. Parker, Benjamin M. Coughlin and Christopher T. Mitchell disclaims beneficial ownership of these shares except to the extent of any pecuniary interest therein.

Reporting Owners
Reporting Owner Name / Address	Relationships
Reporting Owner Name / Address	Director	10% Owner	Officer	Other
Spectrum Equity Investors V L P 333 MIDDLEFIELD ROAD SUITE 200 MENLO PARK, CA 94025		X
Spectrum Equity Associates V LP 333 MIDDLEFIELD ROAD SUITE 200 MENLO PARK, CA 94025		X
Spectrum V Investment Managers Fund LP 333 MIDDLEFIELD ROAD SUITE 200 MENLO PARK, CA 94025		X
SEA V Management LLC 333 MIDDLEFIELD ROAD SUITE 200 MENLO PARK, CA 94025		X
SPECTRUM III INVESTMENT MANAGERS FUND LP 333 MIDDLEFIELD ROAD SUITE 200 MENLO PARK, CA 94025		X
SPECTRUM EQUITY INVESTORS III L P 333 MIDDLEFIELD ROAD SUITE 200 MENLO PARK, CA 94025		X
Spectrum Equity Associates III L P 333 MIDDLEFIELD ROAD SUITE 200 MENLO PARK, CA 94025		X
SEI III Entrepreneurs LLC 333 MIDDLEFIELD ROAD SUITE 200 MENLO PARK, CA 94025		X
SEI III ENTREPRENEURS FUND L P 333 MIDDLEFIELD ROAD SUITE 200 MENLO PARK, CA 94025		X

Signatures
Christopher Brookhart, Attorney-in-Fact		11/4/2009
^** Signature of Reporting Person		Date

Reminder: Report on a separate line for each class of securities beneficially owned directly or indirectly.
*	If the form is filed by more than one reporting person, see Instruction 5(b)(v).
**	Intentional misstatements or omissions of facts constitute Federal Criminal Violations. See 18 U.S.C. 1001 and 15 U.S.C. 78ff(a).
Note:	File three copies of this Form, one of which must be manually signed. If space is insufficient, see Instruction 6 for procedure.
Persons who respond to the collection of information contained in this form are not required to respond unless the form displays a currently valid OMB control number.